Rani

Rani is the main antagonist of the first-half of the third season of The Lion Guard and a supporting protagonist in the second-half. She is the princess (later Queen) of the Tree of Life.

She is more of a Antagonist Hero but she's a de-facto main villain for the first-half. He main goal was to keep Kion and his friends out of the Tree of Life and destroy them. But she later reforms by letting them pass and apologizes for her actions.